Butterflies of India – Annotated Checklist By Paul Van Gasse Kruibeke, Belgium; Email



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Butterflies of India – Annotated Checklist

By Paul Van Gasse (Kruibeke, Belgium; Email: paul.van.gasse@telenet.be), Aug. 2013.


Family Hesperiidae


Subfamily Coeliadinae


  1. Burara oedipodea (Branded Orange Awlet)
    B.o.ataphus: Sri Lanka. NR – Ceylon 17
    B.o.belesis: Kangra to Arunachal, NE India, and Burma to Dawnas (= aegina, athena) – NW Himalayas (Kangra-Kumaon) 11, Sikkim 30, Bhutan 2, Assam 28, Burma (to Dawnas) 9
    B.o.oedipodea: Probably S Burma.
    [Given as Ismene oedipodea in Evans, 1932, and as Bibasis oedipodea in Evans, 1949]

  2. Burara tuckeri (Tucker’s Awlet)
    Burma in Tavoy. VR – Tavoy 1
    [Given as Ismene tuckeri in Evans, 1932, and as Bibasis tuckeri in Evans, 1949]

  3. Burara jaina (Orange Awlet)
    B.j.fergusonii: SW India to N Maharashtra. NR – S India 33
    B.j.jaina: HP (Solan) and Garhwal to Arunachal, NE India, and Burma to Karens. NR (= vasundhara) – NW Himalayas (Dun-Kumaon) 3, Sikkim 18, Assam 37, Burma (Karens) 1
    B.j.margana: Burma in Dawnas. R – Burma (Dawnas) 8
    B.j.astigmata: S Andamans. VR – Andamans 3
    [Given as Ismene jaina in Evans, 1932, and vasundhara was there given as the subspecies ranging from Assam to Karens, with jaina then confined to Mussoorie to Sikkim; given as Bibasis jaina in Evans, 1949]

  4. Burara anadi (Plain Orange Awlet)
    Garhwal to NE India and Burma to Karens. R (= purpurea) – Mussoorie 1, Sikkim 13, Assam 1, Burma (Karens) 5
    [Given as Ismene anadi in Evans, 1932, and as Bibasis anadi in Evans, 1949]

  5. Burara etelka (Great Orange Awlet)
    NE India (Kabaw Valley in Manipur). Burma from Karens S. R – Karens to Mergui 14
    [Given as Ismene etelka in Evans, 1932, and as Bibasis etelka in Evans, 1949]

  6. Burara harisa (Orange Awlet)
    B.h.harisa: C Nepal to NE India and Burma. NR. Recorded from Andamans. (= asambha) – Sikkim 30, Assam 11, Burma 26, Andamans 1
    [Given as Ismene harisa in Evans, 1932, and as Bibasis harisa in Evans, 1949]

  7. Burara vasutana (Green Awlet)
    Kumaon to Arunachal, NE India, and Burma to Dawnas. NR (= rahita, burma) – Nepal 1, Sikkim 25, Assam 27, Burma (Karens, Dawnas) 8
    [Given as Ismene vasutana in Evans, 1932, and as Bibasis vasutana in Evans, 1949]

  8. Burara amara (Small Green Awlet)
    C Nepal to Arunachal, NE India, and Burma to S Shan States. R. Andamans. VR (= pindapatra) – Sikkim 22, Assam 17, Burma (to S Shan States) 14, Andamans 3
    [Given as Ismene amara in Evans, 1932, and as Bibasis amara in Evans, 1949]

  9. Burara gomata (Pale Green Awlet)
    B.g.kanara: SW India to N Kanara and Goa. R – S India 32
    B.g.gomata: Sikkim to Arunachal and NE India. NR – Sikkim 20, Assam 21 – Also listed for Thailand north of Kra.
    B.g.lalita: Burma. (= vajra) – Burma (Karens to Mergui) 5 – Also listed for Thailand south of Kra.
    [Given as Ismene gomata in Evans, 1932, and as Bibasis gomata in Evans, 1949]

  10. Bibasis iluska (Slate Awlet)
    B.i.mahintha: NE India (S Assam and Manipur) and Burma. NR – Manipur 9, Burma (to Ataran) 31
    [Given as Ismene mahintha, Slate Awlet, in Evans, 1932]

  11. Bibasis sena (Orangetail Awl)
    B.s.sena: Sri Lanka. SW India to N Maharashtra and in Madhya Pradesh (Pachmarhi). Pakistan in Murree. Simla to Arunachal, NE India, and Burma to Karens. R. Andamans. VR – Ceylon 8, S India 36, NW Himalayas (Murree, Kumaon) 3, Sikkim 13, Assam 37, Burma (Karens) 2
    B.s.uniformis: Burma from Karens S. NR – Burma (Karens to Ataran) 7

  12. Hasora salanga (Green Awl)
    Burma from Dawnas S. Nicobars (recorded). (= woolletti) – S Burma, Dawnas 4

  13. Hasora anura (Slate Awl)
    H.a.anura: Garhwal to Arunachal, NE India, and Burma to S Shan States. R – Sikkim 10, Khasi Hills 6, Mishmi Hills 2, Naga Hills and Manipur 16, N Burma (Bhamo, Sadon) 7, S Shan States 3

  14. Hasora danda (Smoky Awl)
    NE India (Manipur) and Burma to Karens – Manipur 1, S Shan States 7, Karen Hills 3
    [Described in Evans, 1949; very like anura]

  15. Hasora chromus (Common Banded Awl)
    H.c.chromus: Sri Lanka. All India. Pakistan in Baluchistan (Quetta), Sind (Karachi), and Murree. Nepal. Bangladesh. Burma. S Nicobars. NR. Andamans. VR (= alexis) – Ceylon 16, India (to Baluchistan) 51, Burma 19
    [Given as Hasora alexis, Common Banded Awl, in Evans, 1932]

  16. Hasora taminatus (White-banded Awl)
    H.t.taminatus: Sri Lanka, SW India to S Maharashtra. C (= chromus, Auct., butleri) – Ceylon 20, S India 20
    H.t.bhavara: Kangra. Sikkim to Arunachal, NE India, and Burma to Karens. NR – Kangra 2, Sikkim 14, Assam 23, Burma (to Karens) 4
    H.t.malayana: S Burma. NR (= almea, canostigma, salemana) – S Burma 26
    H.t.andama: Andamans, Car Nicobar. – S Andamans 20, Car Nicobar 4
    H.t.milona: C and S Nicobars. – C Nicobars 10, S Nicobars 2

  17. Hasora schoenherr (Yellow-banded Awl)
    H.s.gaspa: Assam (Nagaland and Manipur) and Burma. – Assam 13, Burma 17
    [Given as Hasora schonherri in Evans, 1932; gaspa was described in 1949]

  18. Hasora mixta (Lesser Awl)
    H.m.prabha: Burma from Dawnas S. R (= lioneli, tyrius, yanuna) – S Burma, Ataran 7
    [Given as Hasora simplicissima lioneli, Simple Awl, in Evans, 1932]

  19. Hasora badra (Common Awl)
    H.b.lanka: Sri Lanka. NR – Ceylon 7
    H.b.badra: SW India to N Maharashtra. C Nepal and adjacent N Bihar to Arunachal, NE India, Bangladesh, and Burma. Andamans, S Nicobars. NR (= certhia, godama, sankarya) – S India 24, Sikkim 20, Assam 15, Burma 21, Andamans 2, Nicobars 1

  20. Hasora vitta (Plain Banded Awl)
    H.v.indica: SW India to N Maharashtra. Sikkim to Arunachal, NE India, and Burma to Dawnas. NR – S India 22, Sikkim 12, Assam 14, Burma (to Karens) 9
    H.v.vitta: S Burma. NR (= chabrona) – S Burma 4
    H.v.manda: Andamans. NR – Andamans 9
    [manda was described as new in 1949]

  21. Hasora khoda (Large Banded Awl)
    H.k.coulteri: NE India (C and S Assam, Meghalaya). Burma in Ataran. Andamans. VR – Assam 7, Ataran 2, Andamans 2

  22. Hasora leucospila (Violet Awl)
    H.l.leucospila: S Burma, Nicobars (recorded). VR (= palinda, matisca, parnia) – Mergui, S Burma 2

  23. Badamia exclamationis (Brown Awl)
    Sri Lanka. All India. Pakistan in Sind (Karachi), Lahore region, Murree, and Chitral. Nepal. Burma. C. Andamans. R (= ericus, ladon, forulus, thymbron) – Ceylon 8, S India 18, Chitral 1, N India 18, Sikkim 16, Assam 14, Burma 12, Andamans 2

  24. Choaspes stigmatus (Branded Awlking)
    C.p.stigmatus: Sikkim to NE India and N Burma. NR – Sikkim 30, Assam 20
    C.p.caudatus: S Burma. NR – Mergui 2
    [Given as Choaspes stigmata, Branded Awlking, in Evans, 1932]

  25. Choaspes benjaminii (Indian Awlking)
    C.b.benjaminii: Sri Lanka, SW India to N Karnataka. NR – Ceylon 17, S India 17
    C.b.japonicus: Kangra to Arunachal and NE India. NR – NW Himalayas (Kangra-Kumaon) 10, Nepal 3, Sikkim 3, Assam 15
    C.b.formosanus: Burma S to Ataran. NR – Burma (Maymyo to Ataran) 20
    [The last two were given as C. benjaminii xanthropogon in Evans, 1932]

  26. Choaspes xanthopogon (Similar Awlking)
    Kashmir to Arunachal and NE India. R (= similis) – NW Himalayas (Kashmir to Kumaon) 6, Nepal 1, Sikkim 7, Assam 23
    [Given as Choaspes similis, Similar Awlking, in Evans, 1932]

  27. Choaspes subcaudatus (Caudate Awlking)
    C.s.crawfurdi: Burma from Karens S. R – Burma (Karens to Mergui) 9

  28. Choaspes furcatus (Hooked Awlking)
    C.h.furcatus: E Nepal to NE India. Burma in Karens. R – Nepal 3, Sikkim 60, Bhutan 1, Assam 9, Burma, Karens 1
    [Given as C. plateni furcata, Hooked Awlking, in Evans, 1932, and as C. hemixanthus furcata in Evans, 1949]


Subfamily Pyrginae



Tribe Celaenorrhini



  1. Capila lidderdali (Lidderdale’s Dawnfly)
    E Nepal. Bhutan. W Arunachal (Dafla Hills). VR – Sikkim or Bhutan (type) 1, Assam (Dafla Hills) 1
    [Given as Orthopaetus lidderdali in Evans, 1932]

  2. Capila phanaeus (Fulvous Dawnfly)
    C.p.fiducia: NE India (Khasi and Lushai Hills) and SE Bangladesh – Khasi Hills 4
    C.p.lalita: NE India (Manipur) to N Burma. R – Manipur 2
    C.p.flora: Burma from N Shan States to Karens. R – N Shan States 6, Thaungyin 1
    C.p.fulva: Burma from Karens to Ataran. R – Karens 14, Rangoon 2, Ataran 6
    C.p.falta: Burma from Ataran to Mergui. R – Ataran 13, Salween 1, Tavoy 7, Mergui 6
    [Given as Orthopaetus phanaeus in Evans, 1932, with lalita and fulva as subspecies; the others were described later]

  3. Capila pennicillatum (Fringed Dawnfly)
    C.p.pennicillatum: E Nepal. NE India (Khasi Hills). VR – Khasi Hills 8
    [Given as Crossiura pennicillatum in Evans, 1932]

  4. Capila pieridoides (White Dawnfly)
    C.p.pieridoides: NE India (N to Margherita region in Assam) and Burma in Dawnas. R – Khasi Hills 31, Manipur 1, Lushai Hills 1, Ataran 1
    [Given as Calliana pieridoides in Evans, 1932]

  5. Capila zennara (Pale Striped Dawnfly)
    Sikkim to Arunachal and NE India. VR – Sikkim 30, Bhutan 2, Manipur 3

  6. Capila jayadeva (Striped Dawnfly)
    E Nepal to Arunachal and NE India. R – Sikkim 12, Assam 28

  7. Capila hainana (Spot Dawnfly)
    C.h.mackwoodi: Burma from N Shan States to Ataran. VR – Burma, Momeit to Ataran 24

  8. Lobocla liliana (Marbled Flat)
    L.l.ignatius: From Murree in Pakistan to Kumaon. R (= casyapa) – NW Himalayas (Murree to Kumaon) 24
    L.l.liliana: C Nepal to Arunachal, NE India, and Burma to Ataran. C – Sikkim 2, Assam 45, N Shan States 2, S Shan States 13, Karen Hills 32, Ataran 2
    L.l.aborica: Arunachal in Abor Hills. VR – Abor Valley 2
    L.l.zesta: Burma in N Shan States. – Momeit 3
    [Given as Achalarus bifasciatus, Marbled Flat, in Evans, 1932]

  9. Celaenorrhinus flavocincta (Bhutan Flat)
    Sikkim, Bhutan. VR – Sikkim 2, Bhutan 3

  10. Celaenorrhinus aspersa (Large Streaked Flat)
    C.a.aspersa: NE India (Nagaland) and Burma to N Shan States. VR (= clitus) – Naga Hills 5, N Burma (Bernardmyo, Momeit) 6

  11. Celaenorrhinus ambareesa (Malabar Flat)
    S India to S Gujarat, Khandesh, Madhya Pradesh, and Chota Nagpur. NR (= hiera) – “Ceylon” (? wrong label) 1, S India 27 DSF, 32 WSF, Pachmarhi 2 DSF, Chota Nagpur 3, “Bengal” 1 WSF (type)

  12. Celaenorrhinus pyrrha (Double Spotted Flat)
    Kumaon to NE India and Burma to S Shan States. NR – Sikkim 7, Bhutan 4, Assam 30, N Burma (Momeit) 1, S Shan States 6

  13. Celaenorrhinus ratna (Tytler’s Multispotted Flat)
    C.r.daphne: Garhwal, Kumaon. R – NW Himalayas (Kumaon) 5
    C.r.tytleri: C Nepal to NE India (Khasi Hills, Manipur). R – Sikkim 33, Assam 35
    [Given as C. tytleri, Tytler’s Multispotted Flat, in Evans, 1932; daphne was described in 1949]

  14. Celaenorrhinus pulomaya (Multispotted Flat)
    C.p.pulomaya: Kangra to Arunachal and NE India (Nagaland). NR (= pila) – NW Himalayas 6, Sikkim 43, Bhutan 3, Assam 37

  15. Celaenorrhinus pero (Mussoorie Spotted Flat)
    C.p.pero: Garhwal, Kumaon. R – Mussoorie and Kumaon 21
    C.p.lucifera: C Nepal, Sikkim, NE India (Nagaland), N Burma. R – Sikkim 1, Naga Hills 3

  16. Celaenorrhinus morena (Pied Flat)
    Sikkim to NE India (C Assam, Nagaland, and Manipur). – Sikkim 2, Manipur and Naga Hills 14 (type from Naga Hills, 6000 ft, Doherty 1889)
    [Described by Evans in 1949; related to plagifera]

  17. Celaenorrhinus plagifera (DeNiceville’s Spotted Flat)
    Sikkim to NE India (Khasi Hills and Nagaland) and N Burma. NR (= moelleri) – Sikkim 28, Bhutan 3, Assam 7, Sadon, N Burma 3
    [Includes C. moelleri, Moeller’s Spotted Flat, an aberration, of Evans, 1932]

  18. Celaenorrhinus sumitra (Moore’s Spotted Flat)
    Sikkim to NE India (Nagaland and Manipur) and Burma to N Shan States. VR – Sikkim 6, Manipur 1, Maymyo, N Shan States 4

  19. Celaenorrhinus patula (Large Spotted Flat)
    C Nepal to Arunachal, NE India, and Burma to Dawnas. R (= pluscula, chinensis) – Sikkim 19, Assam 4, Manipur and Naga Hills 33, Momeit, N Burma 12, S Shan States 11, Karen Hills 6, W Dawnas 1
    [pluscula was considered a synomym of C. plagifera, and chinensis a subspecies of C. leucocera, in Evans, 1932]

  20. Celaenorrhinus leucocera (Common Spotted Flat)
    S India to S Gujarat, Madhya Pradesh, and W Bengal. From Murree in Pakistan to Arunachal, NE India, NE Bangladesh, and Burma; Andamans. C (= arminia, vitruvius) – S India to Centr. Prov. 44, NW Himalayas (Murree to Kumaon) 21, Sikkim 45, Bhutan 5, Assam 23, Burma (to Tavoy) 27, Andamans 19
    [chinensis was considered a subspecies of C. leucocera and recorded from Mishmi Hills in Evans, 1932]

  21. Celaenorrhinus putra (Bengal Spotted Flat)
    C.p.putra: E Nepal to Arunachal and NE India. – Sikkim 6, Bhutan 2, Assam 29
    C.p.sanda: Burma from Karens S. – Burma (Karens to Victoria Point) 20
    [C. putra was given as a synonym of C. leucocera in Evans, 1932; sanda was described in 1941)

  22. Celaenorrhinus munda (Himalayan Spotted Flat)
    C.m.munda: From Murree in Pakistan to Sikkim. R – NW Himalayas (Murree, Simla) 25, Sikkim 8
    C.m.maculicornis: C Nepal to NE India and Burma to Karens. R – Sikkim 2, Assam 25, Burma (to Karen Hills) 11
    [C. maculicornis, Elwes’s Spotted Flat, was considered a separate species in Evans, 1932]

  23. Celaenorrhinus tibetanus (Tibet Flat)
    Arunachal (Mishmi Hills) and Burma in S Chin Hills (Mt Victoria). R – Mt Victoria, Chin Hills, N Burma 1
    [Given as C. tibetana in Evans, 1932]

  24. Celaenorrhinus nigricans (Small-banded Flat)
    C.n.nigricans: Sikkim to Arunachal, NE India, and Burma. NR – Sikkim 2, Bhutan 4, Assam 12, S Shan States 2, Karens 8, Ataran 4, Tavoy 3, Mergui 1

  25. Celaenorrhinus spilothyrus (Black Flat)
    Sri Lanka. C (= infernus) – Ceylon 43

  26. Celaenorrhinus ruficornis (Tamil Spotted Flat)
    C.r.fusca: SW India to S Maharashtra. NR – S India (Palnis, Animalais, Coorg, Nilgiris, N Kanara) 58

  27. Celaenorrhinus badius (Scarce Banded Flat)
    Sikkim to Arunachal (Mishmi Hills) and NE India (Khasi Hills). VR – Sikkim 9, Bhutan 1, Assam 2
    [Given as C. badia in Evans, 1932]

  28. Celaenorrhinus asmara (White-banded Flat)
    C.a.consertus: NE India (Garo and Khasi Hills), SE Bangladesh, and Burma. NR (= cacus, aditta) – Assam 9, N Shan States 2, Karens 18, Rangoon 10, Ataran 5, Tavoy 18, Mergui 13
    [cacus and aditta were considered subspecies, from Rangoon and S Burma, resp., in Evans, 1932]

  29. Celaenorrhinus zea (Swinhoe’s Flat)
    Sikkim to NE India. R – Sikkim 2, Assam 3

  30. Celaenorrhinus dhanada (Himalayan Yellow-banded Flat)
    C.d.dhanada: Garhwal to Bhutan. R – NW Himalayas (Mussoorie, Kumaon) 2, Sikkim 11, Bhutan 6
    C.d.affinis: NE India and Burma to Karens. NR – Assam 19, Burma to Karens 40
    [C. dhanada, Himalayan Yellow-banded Flat, and C. affinis, Burmese Yellow-banded Flat, were considered separate species in Evans, 1932]

  31. Celaenorrhinus andamanicus (Andaman Yellow-banded Flat)
    C.a.hanna: NE India (Jeypore-Dehing Forest in SE Assam) and S Burma from Tavoy S. NR – Tavoy, S Burma 3
    C.a.andamanica: Andamans. VR – Andamans 11
    [Given as C. andamanica in Evans, 1932; hanna was described as new in 1949]

  32. Celaenorrhinus aurivittatus (Dark Yellow-banded Flat)
    C.a.aurivittatus
    : Arunachal (Dibang Valley in Mishmi Hills), NE India, NE Bangladesh, and Burma. Andamans. NR – Assam 21, Burma to Ataran 65, Andamans 1
    C.a.cameroni
    : S Burma in S Mergui. R – S Mergui 1
    [Given as C. aurivittata in Evans, 1932]

  33. Celaenorrhinus ficulnea (Velvet Flat)
    C.f.queda: NE India (Nagaland) and S Burma (= nibana) – Naga Hills 2, S Mergui 4
    [Given as Charmion ficulnea ficulnea, Velvet Flat, in Evans, 1932, where queda was considered a synonym]



Tribe Tagiadini



  1. Tapena thwaitesi (Black Angle)
    T.t.thwaitesi: Sri Lanka, SW India to S Gujarat. S Andhra Pradesh (Nagalapuram Hills). R (= hampsoni) – Ceylon 7, S India (Coorg, Nilgiris, N Kanaras, Khandesh) 71
    T.t.minuscula: NE India (Manipur) and Burma. VR – Manipur 2, Bernardmyo, N Burma 3, N Shan States 2, Ataran 5, Tavoy 1
    [hampsoni was considered the S Indian subspecies in Evans, 1932, with thwaitesi then confined to Sri Lanka]

  2. Darpa hanria (Hairy Angle)
    Garhwal to Arunachal, NE India, and Burma to S Shan States. R – Sikkim 36, Assam 5, S Shan States 1

  3. Darpa striata (Striated Angle)
    D.s.minta: NE India (Manipur) and Burma to Karens. R – Manipur 7, N Shan States 7, Karens 7
    D.s.striata: S Burma from Ataran S. R (= dimidiata) – S Burma (Ataran to Mergui) 20
    [minta was described as new in 1949]

  4. Darpa pteria (Snowy Angle)
    D.p.dealbata: NE India (E Assam and Manipur) and Burma from Karens to Tavoy. R – Assam 1, Tavoy 1

  5. Odina decorata (Zigzag Flat)
    Sikkim to NE India and Burma to Tavoy. R (= bicolor) – Sikkim 2, Assam 5, Burma (to Tavoy) 29
    [Given as O. decoratus in Evans, 1932]

  6. Odina hieroglyphica (Polygon Flat)
    O.h.ortygia: Burma from Ataran S. VR – Ataran 1, Tavoy 1, Mergui 1, Victoria Point 1

  7. Sarangesa purendra (Spotted Small Flat)
    S.p.hopkinsi: Tamilnadu, E Karnataka. R – Madras 14
    S.p.sati: Kutch. NR – Cutch 50
    S.p.pandra: SW India to Gujarat and Madhya Pradesh, and west to Rajasthan. Pakistan in Sind. NR – Ceylon (?) 1, Nilgiris 2, N Kanara 38, Bombay 5, Poona 1, Matheran 1, Mt Abu 4, Pachmarhi, Centr. Prov. 5
    S.p.purendra: Kangra to W Nepal. NR – Kangra 3, Kullu 7, Simla Hills 20, Dun and Garhwal 9, Kumaon 4, Khasi Hills (?) 1
    [Includes S. sati, Tiny Flat, with subspecies hopkinsi, in Evans, 1932; pandra was described in 1949]

  8. Sarangesa dasahara (Common Small Flat)
    S.d.albicilia: Sri Lanka. C (= sezendis) – Ceylon 54
    S.d.davidsoni: SW India to S Gujarat. NR (= hampsoni) – S India to Poona 62
    S.d.adona: Andhra Pradesh and Orissa through Madhya Pradesh to Uttar Pradesh and Rajasthan. C – Pachmarhi 4, Mt Abu 1, Ganjam 2
    S.d.dasahara: Kangra to Arunachal, NE India, NE and SE Bangladesh, and Burma. C (= philippus) – NW Himalayas 10, Behar 1, Sikkim 29, Bhutan 2, Assam 26, Burma to Mergui 47
    S.d.sandra: Andamans. C – Middle Andaman Is 7
    [adona and sandra were described in 1949]

  9. Pseudocoladenia dan (Fulvous Pied Flat)
    P.d.dan: SW India to S Gujarat. S Andhra Pradesh (Nagalapuram Hills). C – S India (Travancore to N Kanara and Mysore) 43
    P.d.fatih: Kangra to Nepal. C – NW Himalayas (Kangra to Nepal) 25
    P.d.fabia: Sikkim to Arunachal, NE India, NE Bangladesh, and Burma to Ataran; flies at lower elevations than P. fatua. – Sikkim 10, Bhutan 4, Assam (700 ft) 32, Burma (to Ataran) 18
    P.d.dhyana: Burma from Dawnas S. C – S Burma (Karens to Mergui) 67
    [Given as Coladenia dan, Fulvous Pied Flat, in Evans, 1932; subspecies fabia was described in 1949 for a population earlier included in fatih]

  10. Pseudocoladenia festa (Naga Pied Flat)
    Sikkim to NE India and Burma to Ataran. C – Sikkim 8, Bhutan 1, Khasi Hills 2, Manipur 2, Naga Hills 13, N Burma (Sadon, Bernardmyo, Bhamo, Momeit, Kambaiti) 19, S Shan States 7, Karens 1, Ataran 1
    [Not in Evans, 1932, but described in 1949]

  11. Pseudocoladenia fatua (Sikkim Pied Flat)
    Sikkim to NE India and N Burma. Occurs in much the same areas as P. festa, but only to N Burma and at lower elevations, and in the plains is replaced by P. dan fabia. – Sikkim 19, Bhutan 1, Lushai 1, Sylhet 1, Khasi Hills 5, Manipur (2-4000 ft) 5, Naga Hills 2, Ruby Mines, N Burma 1

  12. Coladenia indrani (Tricolour Pied Flat)
    C.i.tissa: Sri Lanka. NR (= lankae) – Ceylon 29
    C.i.indra: S India to S Gujarat, Madhy Pradesh, Jharkhand, and lower W Bengal. NR – N Kanara 41, Madras 4, Nilgiris 2, C India, Jabalpur 7, Orissa 2
    C.i.indrani: Kangra to Arunachal and NE India (except Manipur). NR – Kangra 1, Kumaon 2, Sikkim 44, Bhutan 1, Assam 4
    C.i.uposatha: NE India (Manipur) and Burma to Ataran. R (= atarana) – Manipur 2, Burma 35

  13. Coladenia agni (Brown Pied Flat)
    C.a.agni: Sikkim to Arunachal, NE India, and Burma. R – Sikkim 18, Assam 6, Burma (Karens, Ataran, Tavoy) 12

  14. Coladenia agnioides (Elwes’s Pied Flat)
    C Nepal. NE India (Nagaland and Manipur) and N Burma. VR – Manipur 2, Naga Hills 2, Tilin Yaw, N Burma 1

  15. Coladenia buchananii (Great Pied Flat)
    Burma to Karens. – Karen Hills, Burma 1 (type locality: Ruby Mines, N Burma)
    [Considered a synonym of C. laxmi laxmi in Evans, 1932]

  16. Coladenia laxmi (Grey Pied Flat)
    C.l.landa: Sikkim. NE India (C Assam in Panbari Forest, Manipur). VR – Manipur (Sebong) 1
    C.l.laxmi: Burma in Karens. VR – Karens 4
    C.l.sobrina: Burma from S. VR – Ataran to Mergui 5
    [landa was described in 1949]

  17. Satarupa zulla (Tytler’s White Flat)
    S.z.zulla: C Nepal, Sikkim, NE India (Nagaland). VR – Sikkim 10, Naga Hills 4
    [Given as S. majasra zulla, Tytler’s White Flat, in Evans, 1932]

  18. Satarupa gopala (Large White Flat)
    S.g.gopala: E Nepal to NE India. S Burma. NR (= tonkiniana, hainana) – Sikkim 30, Assam 32
    [malaya was given as a subspecies in Evans, 1932, with distribution S Burma]

  19. Satarupa splendens (Splendid White Flat)
    NE India (Nagaland). VR – Naga Hills 2

  20. Seseria dohertyi (Himalayan White Flat)
    S.d.dohertyi: From Murree in Pakistan to Arunachal and NE India. NR – NW Himalayas (Murree to Kumaon) 7, Sikkim 6, Manipur and Naga Hills 3
    [Given as Satarupa dohertyi, Himalayan White Flat, in Evans, 1932]

  21. Seseria sambara (Sikkim White Flat)
    S.s.sambara: Kumaon to Arunachal and NE India. NR (= cosima) – Kumaon 2, Sikkim 35, Bhutan 1, Assam 18, Manipur and Naga Hills (flying with dohertyi) 6
    [Given as Satarupa sambara, Sikkim White Flat, in Evans, 1932]

  22. Seseria strigata (Ringed White Flat)
    Burma from Karens to Dawnas. R – Karen Hills 3, Ataran 8
    [Given as Satarupa kirmana, Ringed White Flat, with strigata as synonym, in Evans, 1932]

  23. Pintara pinwilli (Orange Flat)
    P.p.pinwilli: ? NE India (NE Assam). Burma from Karens S. R – Burma (Karens to Victoria Point) 17
    [Given as Pintara tabrica pinwilli, Orange Flat, in Evans, 1932; Assam occurrence (Margherita, Upper Assam, fide Doherty) doubtful?; tabrica was given with Darjeeling as type locality, but this is very doubtful]

  24. Chamunda chamunda (Olive Flat)
    E Nepal to NE India and Burma to Karens. NR – Sikkim 39, Assam 23, N Shan States 1, S Shan States 2, Karen Hills 4
    [Given as Abraximorpha chamunda, Olive Flat, in Evans, 1932]

  25. Daimio tethys (China Flat)
    D.t.birmana: NE Burma to N Shan States. VR – N Burma (Htaw-gaw, Maymyo) 8

  26. Gerosis sinica (White Yellowbreast Flat)
    G.s.narada: Sikkim to Arunachal, NE India, and Burma to Ataran. NR (= indica) – Sikkim 5 WSF, 27 DSF, Assam 31 WSF, 11 DSF, Ataran, Burma 6 DSF
    [Given as Daimio sinica in Evans, 1932 and 1949; indica was considered a subspecies from Assam in Evans, 1932, with narada then confined to Sikkim]

  27. Gerosis phisara (Dusky Yellowbreast Flat)
    G.p.phisara: Similipal Hills, Orissa. Kumaon to Arunachal, NE India, NE and SE Bangladesh, and Burma. NR (= hamiltonii, kuki, expansa) – Kumaon 1 DSF, Sikkim 17 WSF, 25 DSF, 1 Ab, Assam 28 WSF and Int F, 22 DSF, 1 Ab, Burma (to Tavoy) 1 WSF, 1 DSF, 26 Int F, 2 Ab
    [Given as Daimio phisara in Evans, 1932 and 1949]

  28. Gerosis bhagava (Common Yellowbreast Flat)
    G.b.bhagava: SW India to Goa. Madhya Pradesh (Pachmarhi). Jharkhand. Nepal to Arunachal, NE India, and Burma to Tavoy. NR (= milliana) – S India 4, C India (Pachmarhi) 2, N India (type, from NE Bengal) 1, Nepal 2, Manipur 1, Burma (to Tavoy) 59
    G.b.andamanica: Andamans, Nicobars. R – Andamans 19
    [Given as Daimio bhagava in Evans, 1932 and 1949]

  29. Gerosis limax (Malay Yellowbreast Flat)
    G.l.dirae: Burma from Tavoy S. R (= graya) – S Burma (Tavoy to Mergui) 16
    [Given as Daimio limax in Evans, 1932 and 1949]

  30. Tagiades japetus (Common Snow Flat)
    T.j.obscurus: Sri Lanka. SW India to Goa. NR (= distans) – Ceylon 37, S India (Nilgiris) 13
    T.j.ravi: Madhy Pradesh to Orissa and W Bengal. Garhwal to Arunachal, NE India, Bangladesh, and Burma to Ataran. C (= khasiana, lugens, epicharmus) – Orissa 1, Dehra Dun 1, Calcutta 6, Sikkim Terai 3, Assam 56, Burma (to Ataran) 60
    T.j.atticus: Burma from Tavoy S. C (= utanus, yotissa, rajaghra) – Tavoy to Victoria Point (male Mergui = ravi) 22
    T.j.ravina: Andamans. NR – Andamans 28
    T.j.carnica: Car Nicobar. NR – Car Nicobar 14
    T.j.nankowra: C Nicobars. NR – C Nicobars 26
    T.j.helferi: S Nicobars. NR (= noctis) – Pulo Milo 1, Kondul 4, Little Nicobar 2, S Nicobars 7
    [Given as T. distans, Ceylon Snow Flat, from Sri Lanka, and T. atticus, Common Snow Flat, in Evans, 1932]

  31. Tagiades gana (Suffused Snow Flat)
    T.g.silvia: SW India to S Maharashtra. NR – S India (Travancore to W Ghats and Chota Nagpur) 52
    T.g.athos: Orissa, Jharkhand, W Bengal, and C Nepal to Arunachal, NE India, and NE Bangladesh. NR – Orissa 1, Nepal 2, Sikkim 4, Assam 27
    T.g.meetana: Burma. NR (= patimoka) – Burma (Pegu Yoma and Karens to Mergui) 46
    T.g.gana: S Burma (Mergui). (= menanto, avala, paceka, mahinda, perakana, javanus) – S Burma (Mergui to Victoria Point) 11
    T.g.alica: Andamans. NR – Andamans 38, C Nicobars 2
    [Given as T. obscurus, Suffused Snow Flat, in Evans, 1932; silvia was described by Evans in 1934, S India being included in the range of athos before that]

  32. Tagiades parra (Large Snow Flat)
    T.p.gala: C Nepal to NE India and Burma. R – Sikkim 42, Assam 6, Burma (to Mergui) 13
    [Given as T. gana, Large Snow Flat, in Evans, 1932; gala was described as new in 1949]

  33. Tagiades lavatus (Scarce Snow Flat)
    S Burma from Tavoy S (= albovittata) – S Burma (Tavoy, Victoria Point) 2
    [Given as T. lavata in Evans, 1932 and 1949]

  34. Tagiades calligana (Malayan Snow Flat)
    Malaysian region north to Peninsular Thailand. (= yapatha) – Naga Hills (?) 1

  35. Tagiades toba (Small Snow Flat)
    T.t.toba: S Burma in Mergui. R – S Burma (Mergui) 6

  36. Tagiades litigiosa (Water Snow Flat)
    T.l.ceylonica: Sri Lanka. NR – Ceylon 28 WSF, 11 DSF
    T.l.litigiosa: S India to N Maharashtra (Tamhini region). S Andhra Pradesh (Nagalapuram Hills). Jharkhand. Simla to Arunachal, NE India, NE and SE Bangladesh, and Burma to Tavoy. NR (= vajuna, eson, kotoshona) – S India 42, NW Himalayas (?) 2, Sikkim 31, Assam 30, Burma (to Ataran) 26
    T.l.andamanica: Andamans. NR (= major, DSF) – Andamans 7 WSF, 4 DSF

  37. Tagiades ultra (Ultra Snow Flat)
    Burma from Tavoy S. NR – S Burma (Tavoy, Mergui) 9
    [Given as a race of T. litigiosa in Evans, 1932]

  38. Tagiades menaka (Spotted Snow Flat)
    T.m.menaka: Kashmir to NE India and Burma to Ataran. C (= vulturna, gavina) – NW Himalayas (Kullu to Kumaon) 30, Sikkim 47, Bhutan 3, Assam 44, Burma (to Ataran) 13

  39. Tagiades cohaerens (Evans’s Snow Flat)
    T.c.cynthia: Murree to NE India and Burma to Karens. (= pintra, WSF) – NW Himalayas (Murree) 1 WSF, Nepal 1 WSF, Sikkim 1 DSF, Assam 6 DSF, 7 WSF, S Shan States 3 DSF, 5 WSF, Karen Hills 15 DSF, 1 WSF
    [cynthia and pintra were described by Evans in 1934; cohaerens, described from Taiwan, was considered a synonym of T. litigiosa litigiosa in Evans, 1932]

  40. Mooreana trichoneura (Yellow Flat)
    M.t.pralaya: Kumaon to Arunachal, NE India, and Burma to Karens. NR (= pellita) – Kumaon 1, Sikkim 18, Assam 23, N Burma (Ruby Mines) 1, S Shan States 1, Karens 1
    M.t.trichoneura: Burma from Karens S. NR – Karens 8, Ataran 19, Tavoy 5, Mergui 5

  41. Abraximorpha davidii (Chequered Flat)
    A.d.esta: N Burma to S Shan States. R – N Burma (Sadon) 1, S Shan States 2
    [Comment: Sometimes treated as a separate species, A. esta]

  42. Ctenoptilum vasava (Tawny Angle)
    C.v.vasava: Garhwal to Arunachal, NE India, and Burma. NR – Sikkim 23, Assam 24, Burma (to Tavoy) 28

  43. Ctenoptilum multiguttatum (Multispot Angle)
    NE India (Manipur) and Burma to Ataran. R – Manipur 12, Burma (to Ataran) 37
    [Given as C. multiguttata in Evans, 1932 and 1949]

  44. Odontoptilum angulatum (Chestnut Angle)
    O.a.angulatum: Sri Lanka. SW India to N Maharashtra (Tamhini region). Kangra to Arunachal, NE India, NE and SE Bangladesh, and Burma. NR (= sura, sumatrana, mahabini) – Ceylon 2, S India 27, NW Himalayas (Kangra, Kullu) 4, Nepal 1, Sikkim 19, Bhutan 3, Assam 27, Burma 34
    [Given as O. angulata in Evans, 1932 and 1949]

  45. Odontoptilum pygela (Banded Angle)
    O.p.pygela: Burma from Karens S. NR (= ragupta) – Burma (Karens to Mergui) 18

  46. Caprona ransonnetii (Golden Angle)
    C.r.ransonnetii: Sri Lanka. R. – Ceylon 43 WSF
    C.r.potiphera: S India to S Gujarat, Madhya Pradesh, Chhattisgarh, Orissa, and Jharkhand. NR (= taylorii, DSF, pexadora) – S India to Bombay 51 WSF, 31 DSF, Pachmarhi 1 WSF, Orissa 1 WSF, 3 DSF

  47. Caprona agama (Spotted Angle)
    C.a.agama
    : From S India very locally north to Maharashtra, Madhya Pradesh, Orissa, and Jharkhand. Pakistan in N Punjab. Garhwal to NE India and Burma to S Shan States. NR (= syrichthus, erosula, danae, saraya, pelias, pelligera, mettasuta) – WSF: S India (Palnis, Madura) 3, C India (Jabalpur, Chota Nagpur) 2, Orissa 3, Bhutan 2, Assam 4, Manipur 16, Burma 46 – DSF: W Punjab (near Rawalpindi) 1, Nepal 2, Khasi Hills 7, Manipur 7, N and S Shan States 17

  48. Caprona alida (Yellow-spotted Angle)
    C.a.lanka
    : Sri Lanka. R – Ceylon 5
    C.a.vespa
    : From S India very locally north to Madhya Pradesh and Orissa. R – Nilgiris 5, Coorg 1, Jabalpur, Centr. Prov. 1, Orissa 1
    C.a.yerburyi
    : Pakistan in N Punjab (Attock). Haryana (Kalka). Garhwal (Dehra Dun). W and C Nepal terai and adjacent Bihar (Jamalpur). – Chittar Pahar (near Rawalpindi, Punjab) 6, Jamalpur (Behar) 1, Dehra Dun 1, Nepal 1
    C.a.alida
    : Arunachal, NE India (Khasi Hills, Manipur), and Burma to Dawnas. NR (= elwesii, parvipunctata) – DSF (alida): Kalewa, Chindwin 1, “Lower Burma” 1 – WSF (elwesii): Manipur 2, Chin Hills 3, Bhamo 1, Ruby Mines 5, N Shan States 5
    [These were included in C. ransonnetii, Golden Angle, in Evans, 1932; lanka was considered the Sri Lanka and Nilgiri subspecies, vespa and yerburyi were described in 1949, and alida was also considered a subspecies]

  49. Erynnis pathan (Chitral Inky Skipper)
    E.p.max: Pakistan in N Baluchistan and Waziristan. NR – Baluchistan, Pishin Valley (Quetta to Chaman and Zhob watershed) 29
    E.p.pathan: Pakistan in Chitral and Swat. NR – Shingarh, near Fort Sandeman, Zhob Valley 2, Razmak, NWFP 1, Chitral 16, Bashahr, NW Himalayas 3
    [Described by Evans in 1949, as E. marloyi max and E. marloyi pathan]

  50. Gomalia elma (African Marbled Skipper)
    G.e.albofasciata: Sri Lanka. S India to N Maharashtra and NW Andhra Pradesh (Hyderabad region). Kangra and Kullu. Pakistan in S Sind and C Baluchistan, and recorded N Punjab (Rawalpindi). R (= litoralis) – Baluchistan 3, Kullu 2, Karachi 1, S India to Poona 82, Ceylon 2
    [litoralis, from Sind and Baluchistan, was considered a subspecies in Evans, 1932)

  51. Carcharodus alceae (Plain Marbled Skipper)
    C.a.swinhoei: Pakistan in N Baluchistan. C – Baluchistan 70
    C.a.gooraisa
    : From Chitral in Pakistan to Garhwal. NR – Chitral 35, Kashmir 28, Murree Hills 40, Simla 2, Mussoorie 1
    [gooraisa was described as new in 1949; that population was included in C.a.alceae before that]

  52. Carcharodus floccifera (Tufted Marbled Skipper)
    C.f.dravira
    : Pakistan in N Baluchistan. NR. Chitral in Pakistan and Kashmir. R (= balucha, dentatus) – Baluchistan (May-Sept) 46, Chitral 16, Kashmir 12
    [Given as C. altheae, Tufted Marbled Skipper, in Evans, 1932, with subspecies dravira, from Chitral and Kashmir, and balucha, from Baluchistan]
    [Comment: Sometimes treated as a separate species, C. dravira]



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